Friday, January 19, 2018

Ontario: Saddlebags in July

The 2015 Ontario dragonfly-watching season continued into July, and on the 11th Ryan and I paid a visit to the Fletcher Creek Ecological Preserve near Puslinch.

Fletcher Creek is a rehabilitated fen wetland - a former quarry that has now become an example of one of Ontario’s rarer habitat types. It is an excellent spot for dragonflies (which is why, of course, we were there).

Dragonflies and damselflies vary in the timing and season of their adult flight period, so even the space of a few weeks made a considerable difference as to which species we were likely to see. 

Ebony Jewelwing (Calopteryx maculata)
Some of the species we came across, of course, were the same we had seen nearby a month earlier (see my previous post).   They included the lovely Ebony Jewelwing (Calopteryx maculata)...

Eastern Forktail (Ischnura verticalis)
Eastern Forktail (Ischnura verticalis)
Eastern Forktail (Ischnura verticalis)
...and the ubiquitous Eastern Forktail (Ischnura verticalis), which flies well into the autumn.   Perched male Eastern Forktails frequently wave their abdomens back and forth, presumably to enhance the display effect of its bright blue tip.


Eastern Forktail (Ischnura verticalis)
This is an Eastern Forktail too - an immature female... 

Eastern Forktail (Ischnura verticalis)
...and an even more immature, or teneral, individual, only recently emerged and still to gain adult colouration.

Bluet (Enallagma sp)
Bluet (Enallagma sp)
Bluets, as usual, were about, mating, and largely (to me) unidentifiable.

Marsh-type Bluet (Enallagma cf ebrium)
However, this one could be a Marsh Bluet (Enallagma ebrium), based on the pattern of blue and black on the abdominal segments. 

Boreal Bluet (Enallagma aspersum)
Boreal Bluet (Enallagma aspersum
We had the good fortune to run across another dragonfly enthusiast, who was far better informed than I about identification matters. He pointed out that these particular individuals (and others of their type) were Boreal Bluets (Enallagma aspersum). I believe this was based on prior knowledge of what was in the area, as these insects are almost impossible to identify definitively without a close examination of the male’s genitalia. 


Violet Dancer (Argia fumipennis violacea)
Violet Dancer (Argia fumipennis violacea)
Violet Dancer (Argia fumipennis violacea)
The Violet Dancer (Argia fumipennis violacea) is much easier to identify. No other damselfly in Ontario is this colour. They have a particular penchant for perching on rocks, pieces of wood or gravel pathways.


Violet Dancer (Argia fumipennis violacea)
This is the local representative among the  three well-marked subspecies of the Variable Dancer (Argia fumipennis); the other two look quite different

Four-spotted Skimmer (Libellula quadrimaculata)
Among the dragonflies at Fletcher Creek we found a number of species we had missed on our June excursions. They included the Four-spotted Skimmer (Libellula quadrimaculata), a widespread species that also occurs in Europe. 

Widow Skimmer (Libellula luctuosa)
It’s cousin the Widow Skimmer (Libellula luctuosa) is one of our handsomest dragonflies. This is a young male, with the distinctive wing pattern of the adult but without the heavy bluish-white pruinosity, or powdery covering, concealing his yellow-striped abdomen. 

Dot-tailed Whiteface (Leucorrhinia intacta)
Dot-tailed Whiteface (Leucorrhinia intacta)
Whitefaces (Leucorrhinia spp) are smallish dragonflies with a distinctly northern distribution in both the Old and New Worlds. The Dot-tailed Whiteface (Leucorrhinia intacta) is probably the commonest of the seven North American species, and is most easily identified by the yellow spot on the top of its seventh abdominal segment (counting, as odonatologists do, from the front to the back).


Frosted Whiteface (Leucorrhinia frigida)
Whitefaces are variable and tricky to identify, but I believe this one is a Frosted Whiteface (Leucorrhinia frigida) based on the powdery 'frosting', or pruinosity, on the upper side of ts abdomen (as opposed to on the underside, a in the Dot-tailed Whiteface).


Black Saddlebags (Tramea lacerata)Black Saddlebags (Tramea lacerata)
Black Saddlebags (Tramea lacerata)
The prize of the day, for me, was my first decent sighting of a Black Saddlebags (Tramea lacerata) at the ege of one of the larger ponds.  This is not a rare species, but one that I had been unable to get more than a glimpse of before.  Like other Tramea species, it has a dark patch on the base of the hindwing (the so-called saddlebag).


Black Saddlebags (Tramea lacerata)
Black Saddlebags are not only wide-ranging (they have even reached Hawaii) but they are highly migratory.  Numbers have been seen massing on the shores of Lake Ontario in autumn along with Ontario's other major dragonfly migrant, the Common Green Darner (Anax junius), preparing to strike out for the south.  Like the better-known Monarch Butterfly (Danaus plexippus), the dragonflies that leave our shores will not return; it is their offspring that will make the spring journey northward.


Eastern Comma (Polygonia comma)Eastern Comma (Polygonia comma)
Though Ryan and I were on an odonatological expedition, we did not ignore the other creatures on the reserve.  We watched butterflies too, including this Eastern Comma (Polygonia comma)...


Baltimore Checkerspot (Euphydryas phaeton)
Baltimore Checkerspot (Euphydryas phaeton)
Baltimore Checkerspot (Euphydryas phaeton)
... and a particularly lovely little insect, the Baltimore Checkerspot (Euphydryas phaeton).  There is nothing else quite like it in eastern North America, though there are a number of other checkerspots, members of the nymphalid subfamily Melitaeinae, in the west.

Pickerel Frog (Lithobates palustris)
The Pickerel Frog (Lithobates palustris) resembles the more common leopard frogs, but is distinguished by its greyish-brown ground colour and the squarish, rather than round, spots on its back.  It gets its name, I am sorry to say, from the tendency of pickerel fishers to use it for bait.


Northern Water Snake (Nerodia sipedon) Northern Water Snake (Nerodia sipedon)
Northern Water Snake (Nerodia sipedon)
Undoubtedly some of the Pickerel Frogs that are not devoured by their fishy namesakes will end up in the stomachs of Northern Water Snakes (Nerodia sipedon sipedon).  This is one of our commonest snakes, and (for the benefit of any Ontario readers I may have) it is not a Water Moccasin (Agkistrodon piscivorus) (which does not occur in Ontario) and is not venemous.


Rufous-sided Towhee (Pipilo erythrophthalmus)Rufous-sided Towhee (Pipilo erythrophthalmus)
Finally, just to show that even though it was July we had not given up birding altogether, here is a Rufous-sided Towhee (Pipilo erythrophthalmus) singing from the top of a shrub.  Ryan and I are all-around naturalists, we are.

Ontario: Baskettails in June

As May turns into June and spring advances towards summer, birders in southern Ontario have less and less to do. The migrant warblers that keep us on the chase in May have, except for a few resident species, moved on to their northern breeding grounds, and many of the birds that stay behind, busy raising their broods, have started to fall silent and retreat to the now fully-leafed-out cover of the woodland canopy. Birders, in short, need something to keep them occupied, and for some of us this means turning to the emerging crop of damselflies and dragonflies. June and July are the peak damselfly- and dragonfly-watching months here, and with Ryan and his brothers still with us in Canada I could justify an insect-hunting expedition or two in June 2015 as part of my schedule of grandpa-grandson bonding experiences (sometimes, admittedly, under some protest from Ryan). 

The best dragonfly spots near Toronto and Mississauga, so I had been told, are actually well west of the city in the vicinity of Cambridge and Puslinch. June 6, therefore, found us exploring the Shade's Mills Conservation Area on the Grand River near Cambridge.  Shade’s Mills turned out to be quite an attractive place, a combination of hardwood forest...

...and waterside that proved to be attractive to both insects...

...and to small boys who enjoy poking about among the reeds. 

Northern Leopard Frog (Lithobates pipiens)
Northern Leopard Frog (Lithobates pipiens)
After all, you never know what a few judicious pokes might turn up: perhaps, with luck, a frog or two (in this case, a Northern Leopard Frog (Lithobates pipiens) hunkering down among the grass stems). 

Yellow Iris (Iris pseudacorus)
While Ryan checked for frogs, I concentrated on attractive waterside plants like this blooming Yellow Iris (Iris pseudacorus)...

Dark Fishfly (Nigronia fasciata)
...and the occasional insect. This is a Dark Fishfly (Nigronia fasciata). 

Canada Anemone (Anemone canadensis)
I also paid attention to woodland wildflowers. This is Canada Anemone (Anemone canadensis). 

Six-spotted Tiger Beetle (Cicindela sexguttata)
Six-spotted Tiger Beetles (Cicindela sexguttata) apparently take a long time mating...

Six-spotted Tiger Beetle (Cicindela sexguttata)
Six-spotted Tiger Beetle (Cicindela sexguttata)
...or at least they spend a fair bit of time wandering around locked together, the male clinging to the female like a frog in amplexus. 

Forest Tent Caterpillar (Malacosoma disstria)
It’s not easy to find a Forest Tent Caterpillar (Malacosoma disstria) on its own. It’s much more common to find masses of these colonial caterpillars, hiding within their silken enclosures from predators such as cuckoos (my old mentor, the late Professor J. Murray Speirs, used to refer to tent caterpillar colonies as “cuckoo food”).

Delaware Skipper (Anatrytone logan)
Delaware Skipper (Anatrytone logan)
Delaware Skipper (Anatrytone logan)
The Delaware Skipper (Anatrytone logan) is an uncommon butterfly in Canada, but it has been spreading in southern Ontario in recent years. 

Red Admiral (Vanessa atalanta)
Red Admirals (Vanessa atalanta), on the other hand, are among our commonest butterflies. 


Eastern or Canadian Tiger Swallowtail (Papilio glaucus or canadensis)
Eastern or Canadian Tiger Swallowtail (Papilio glaucus or canadensis)
Southern Ontario sits on the border between the ranges of two spectacular, but very similar, butterflies, the Eastern and Canadian Tiger Swallowtails (Papilio glaucus and P. canadensis). Unfortunately these photos don’t make it clear (at least to me) which of the two this is. The best way to tell is to check whether the spots in a row on the underside of the forewing edge butt up against each other (Canadian) or not (Eastern). 

Giant Swallowtail (Papilio cresphontes)
In recent years a warming climate has allowed Giant Swallowtails (Papilio cresphontes) to stage an invasion of much of southern Ontario. Previously confined, at least in our province, to the Carolinian lowlands of the southwest, Giants have begun appearing over a far wider area. 

Giant Swallowtail (Papilio cresphontes)
Giant Swallowtail (Papilio cresphontes)
Their distribution is probably limited here by the availability of larval food plants. The orange groves that feed broods of Giant Swallowtails in places like Florida are not available here, and their caterpillars have to make do with related substitutes such as Hoptree (Ptelea trifoliata), a relatively rare plant in Ontario, and Prickly-Ash (Zanthoxylum americanum). 

Ebony Jewelwing (Calopteryx maculata)
As for the objects of our expedition, Shade’s Mills proved to be a good spot for both of Ontario’s brilliant jewelwing damselflies (Calopteryx spp). 

Ebony Jewelwing (Calopteryx maculata)
The Ebony Jewelwing (Calopteryx maculata), as usual the commoner of the two, must be one of the most elegant of its kind. The male, his broad black wings draped over his deeply iridescent body, looks downright aristocratic. 

Ebony Jewelwing (Calopteryx maculata)
The female, easily recognized by the white stigmas in her wings, is almost as spectacular. 

River Jewelwing (Calopteryx aequabilis)
The other species, the River Jewelwing (Calopteryx aequabilis), is not quite a match for the Ebony. Its wings are not as broad, and the black is confined to the wingtips  it is, of course, a very handsome insect all the same.  

Ebony Jewelwing (Calopteryx maculata)
Ebony Jewelwing (Calopteryx maculata)
The females of the two species are a bit harder to tell apart, but the female River Jewelwing (lower photograph) does have a duller version of the male’s wing pattern.  Both sexes have, as I noted for the male, narrower wings than in the Ebony Jewelwing (upper photograph). 

Ebony Jewelwing (Calopteryx maculata)
Ebony Jewelwing (Calopteryx maculata)
Ebony Jewelwing (Calopteryx maculata)
We found both jewelwings posing, in typical fashion, on the upper surfaces of leaves along trails through the forest.  These are Ebony Jewelwings.

Eastern Forktail (Ischnura verticalis)
Eastern Forktail (Ischnura verticalis)
Eastern Forktail (Ischnura verticalis)
In addition to both species of local jewelwing, we also found numbers of Eastern Forktails (Ischnura verticalis). The upper insect is an  adult male. Young female Forktails are heteromorphic (that is, they coe in more than one colour), and may be orange-and-black (middle) before settling down to the mostly blue colour of the adult (bottom).

Fragile Forktail (Ischnura posita)
I at first took this for the smaller and daintier Fragile Forktail (Ischnura posita), usually best identified by the ‘exclamation mark’ on the thorax, but the Fragile Forktail never has blue on the tip of the abdomen. This is, then, simply another male Eastern Forktail, but one with an interrupted shoulder stripe – something that turns up not infrequently.

Bluet (Enallagma sp)
Bluet (Enallagma sp)
The commonest of our damselflies are probably the Bluets (Enallagma spp). Finding them is easy, but identifying them - short of catching them, which I don’t like to do, and examining their genitalia with a hand lens - can be difficult to impossible. 

Bluet (Enallagma sp)
Bluet (Enallagma sp)
So these must go down as simply as unidentified bluets, alas. 

Common Whitetail (Plathemis lydia)
Common Whitetail (Plathemis lydia)
Common Whitetail (Plathemis lydia)
Of the several dragonfly families recorded from Ontario one - the Libellulidae, or Skimmers - normally accounts for most of the dragonflies one is likely to see and photograph, including these Common Whitetails (Plathemis lydia). The upper photograph shows an adult male; the other two are females. 

Lancet Clubtail (Phanogomphus exilis)
Lancet Clubtail (Phanogomphus exilis)
Other than that, the family I encounter most often is the Gomphidae, or Clubtails, easily identified by their damselfly-like eyes that do not, as in most other dragonflies, meet across the top of the head. 

Lancet Clubtail (Phanogomphus exilis)Lancet Clubtail (Phanogomphus exilis)
I found two common species sitting (as they typically do) on the open ground. These are Lancet Clubtails (Phanogomphus exilis).  

This is the very similar Ashy Clubtail (Phanogomphus lividus). Alert readers will notice that when I have shown these two species in previous posts I have listed them in the genus Gomphus. Gomphus, though, has now been split into a number of genera according to the regularly-updated World Odonata Listand who am I to argue? 

Ashy Clubtail (Gomphus lividus)
Anyway, the split is based on a 2017 study that was the first to examine North American Gomphidae using molecular evidence. According to the study, as it turns out, none of the North American dragonflies included in the genus Gomphus actually belong there. Who knew?

Common Baskettail (Epitheca cynosura)
With the exception of the Aeshnidae, or Darners, I had, so far, had miserable luck (or simply lousy locating skills) with Ontario’s other dragonfly families. This was particularly galling for the Corduliidae, or Emeralds. Ontario actually has lots of corduliids, but my experience with them had been limited to brief, distant looks and poor photographs. My luck changed at Shade’s Mills, at least with respect to the commonest (and, I suppose, easiest to see) genus of corduliids, the baskettails (Epitheca spp.)

Common Baskettail (Epitheca cynosura)
Baskettails seemed to be pretty much everywhere at Shades Mills, at least in more open areas. They proved to be distinctive creatures, with their colorful eyes, hairy bodies and habit of clinging to vertical perches while holding their bodies diagonally or even horizontally - a position that, for a human gymnast, would be challenging in the extreme, but seems normal for this dragonfly family. 

Spiny Baskettail (Epitheca spinigera)
Spiny Baskettail (Epitheca spinigera)
Identifying Baskettails to genus is comparatively easy (though, in my subsequent encounters with them, I have misidentified then more than once), but sorting out the species can be a lot trickier. The key, again, is the shape of the genitalia. In the Spiny Baskettail (Epitheca spinigera) there is, indeed, a tiny spine projecting down from the underside of the upper male claspers (or cerci). You can just see it in these photographs. 

Common Baskettail (Epitheca cynosura)
Common Baskettail (Epitheca cynosura)
Common Baskettail (Epitheca cynosura)
The spine is lacking in the Common Baskettail (Epitheca cynosura), a distinction I did not notice until I examined my photographs at home. Just to make things slightly easier, the Common is more likely than the Spiny to have a patch of black at the base of the hindwing. 

Wood Frog (Lithobates sylvaticus
For a final image from Shade’s Mills, here is a Wood Frog (Lithobates sylvaticus) that Ryan and I found, appropriately, in the woods. Wood Frogs are northern, highly cold-tolerant animals, and unlike other ranid frogs are more creatures of the forest floor than the water’s edge. 

Common Green Darner (Anax junius)
I mentioned the Darner family (Aeshnidae) earlier in this post, but didn’t include any photos. Darners are difficult to photograph because, unlike skimmers, clubtails and emeralds, they rarely perch during their hunting sessions. I was therefore very pleased, during a brief excursion on June 21, to come across a mating pair of Common Green Darners (Anax junius) conveniently perched at close range. Here is the result.